Nikon updated their workhorse 70-200 f/2.8 lens with lots of new tech! The new E designation means it employs an electronic diaphragm which syncs with the camera's shutter to give consistent, accurate exposures during high drive modes. No more physical aperture lever! Nikon promises it has gotten rid of the previous lens's focus breathing problems at longer focal lengths, sure to please videographers. New elements are coated in new fancy coatings with such fancy terms as Fluorite, High Refractive Index, and Nano Crystal Coatings. All these should add up to an increase in sharpness and a decrease in glare, even in backlit conditions. Nikon also added 4 focus function buttons to the lens body, similar to its high end super-telephoto lenses. These buttons are able to engage autofocus or AF Lock on certain bodies. The zoom and focus rings have been switched; the zoom ring is now furthest from the camera and the focus ring is now closest to the camera. Nikon says this is to make zooming more comfortable and balanced. The minimum focusing distance has decreased from about 4.5 feet down to about 3.5 feet. Vibration Reduction (VR) should give you about 4 stops of relief when shooting handheld.